TYPE: Costa Rica. Puntarenas: Monteverde Reserve, near Alajuela boundary, Cerros Centinelas, elev. 1550-1600 m, 10.18N 84.47W, terrestrial in springy soil in cloud forest along ridge, Sendero Brillante; flowers with dull reddish tinge, 19 Aug 1984, M. H. Grayum & P. Sleeper 3842 (holotype, MO; illustration of type).